Dan received a BA degree in chemistry at Colby College in 1968. He received his Ph. D. in organic chemistry from Penn State University in 1974. In 2015, Dan retired after 41 years of undergraduate teaching and research at Moravian University, as well as at Colby, Barnard, Skidmore, Kenyon and Oberlin Colleges. His research involved mechanisms of enzyme reactions. Dan has used personally written learning cycle-based activities to teach all of his courses since 1980 and has facilitated POGIL workshops across the USA and in India.